Monofenestrata

Clade of breviquartossan pterosaurs

Monofenestratans
Temporal range:
Middle Jurassic – Late Cretaceous, 166–66 Ma
PreꞒ
O
S
D
C
P
T
J
K
Pg
N
Fossil specimen of Darwinopterus modularis
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Clade: Pterodactylomorpha
Clade: Monofenestrata
et al., 2010
Subgroups

Monofenestrata is an unranked group of pterosaurs that includes the family Wukongopteridae and the suborder Pterodactyloidea.[2]

The clade Monofenestrata was in 2009/2010 defined as the group consisting of Pterodactylus and all species sharing with Pterodactylus the synapomorphy of an external nostril confluent with the antorbital fenestra, the major skull opening on the side of the snout. The name is derived from Greek monos, "single", and Latin fenestra, "window". The concept was inspired by the discovery of Darwinopterus, a species combining a pterodactyloid-type skull with a more basal build of the remainder of the body.[2] The Darwinoptera, a primitive subgroup of monofenestratans showing this transitional anatomy, was also named for Darwinopterus and defined as all descendants of its common ancestor with Pterorhynchus.[3]

The earliest known monofenestratan fossils have been found in the Stonesfield Slate formation of the United Kingdom, which dates to the Bathonian stage of the Middle Jurassic, dated to about 166 million years ago. Identified elements include cervical vertebrae, fourth metacarpals and a possible pterodactyloid synsacrum. [4]

Below is a cladogram showing the results of a phylogenetic analysis presented by Andres, Clark & Xu, 2014. This study found the two traditional groupings of ctenochasmatoids and kin as an early branching group (represented as the group Archaeopterodactyloidea), with all other pterodactyloids grouped into the Eupterodactyloidea.[3]

References

  1. ^ a b Martin-Silverstone, Elizabeth; Unwin, David M.; Cuff, Andrew R.; Brown, Emily E.; Allington-Jones, Lu; Barrett, Paul M. (2024-02-05). "A new pterosaur from the Middle Jurassic of Skye, Scotland and the early diversification of flying reptile". Journal of Vertebrate Paleontology. doi:10.1080/02724634.2023.2298741. ISSN 0272-4634.
  2. ^ a b Lü, J.; Unwin, D.M.; Jin, X.; Liu, Y.; Ji, Q. (2010). "Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull". Proceedings of the Royal Society B. 277 (1680): 383–389. doi:10.1098/rspb.2009.1603. PMC 2842655. PMID 19828548.
  3. ^ a b Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24 (9): 1011–6. doi:10.1016/j.cub.2014.03.030. PMID 24768054.
  4. ^ Michael O’Sullivan; David M. Martill (2018). "Pterosauria of the Great Oolite Group (Bathonian, Middle Jurassic) of Oxfordshire and Gloucestershire, England". Acta Palaeontologica Polonica. Online edition. doi:10.4202/app.00490.2018.
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Avemetatarsalia
    • see Avemetatarsalia
Pterosauria
    • see below↓
Preondactylia
Caviramidae?
Austriadraconidae
Eudimorphodontoidea
Raeticodactylidae
Eudimorphodontidae
Eudimorphodontinae
Dimorphodontidae
Campylognathoididae
Rhamphorhynchidae
Rhamphorhynchinae
Rhamphorhynchae
Angustinaripterini
Rhamphorhynchini
Digibrevisauria?
Scaphognathidae?
Pterodactylomorpha
    • see below↓
Campylognathoides liasicus

Scaphognathus crassirostris

Dorygnathus banthensis
Monofenestrata
Darwinoptera
Wukongopteridae
Wukongopterinae
Anurognathidae
Anurognathinae
Batrachognathinae
Pterodactyloidea
Lophocratia
    • see below↓
Jeholopterus ninchengensis Kryptodrakon progenitor
Germanodactylidae
Gallodactylidae
Aurorazhdarchia
Aurorazhdarchidae
Moganopterinae?
Gnathosaurinae
Ctenochasmatinae
Pterodaustrini
Eupterodactyloidea
Ornithocheiroidea
    • see below↓
Pterodactylus antiquus Plataleorhynchus streptorophorodon
Dsungaripteridae
Dsungaripterinae
Thalassodrominae?
Caupedactylia?
Eutapejaria
Sinopterinae
Tapejarinae
Tapejarini
Alanqidae?
Neopterodactyloidea
Chaoyangopteridae
Chaoyangopterinae
Azhdarchiformes
Azhdarchidae
Azhdarchinae
Quetzalcoatlinae
Pteranodontoidea
    • see below↓
Bakonydraco galaczi

Tupandactylus imperator

Quetzalcoatlus
Pteranodontia
Pteranodontidae
Nyctosauromorpha
Aponyctosauria
Nyctosauridae
Lonchodectidae
Lonchodraconidae
Istiodactyliformes
Mimodactylidae
Istiodactylidae
Istiodactylinae
Ornithocheiriformes
Boreopteridae
Ornithocheiridae
Ornithocheirinae
Targaryendraconia?
Cimoliopteridae
Targaryendraconidae
Hamipteridae?
Anhangueridae
Tropeognathinae?
Coloborhynchinae?
Anhanguerinae
Pteranodon longiceps

Nyctosaurus gracilis

Ludodactylus sibbicki
Taxon identifiers
Monofenestrata